5. When conservators examined medieval manuscripts, they once focused primarily on the words written by scribes and the illustrations added by artists. More recently, however, researchers have also studied stains, repairs, fingerprints, and worn page edges. These physical traces can suggest which pages were handled most often, whether a manuscript was used in a classroom or religious setting, and how later owners modified the book. As a result, scholars increasingly treat manuscripts not only as texts but also as objects with histories of use.
Which choice best states what can reasonably be inferred from the text?